Esenbeckia berlandieri subsp. acapulcensis (Rose) Kaastra
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Authority
Kaastra, Roelof C. 1982. A monograph of the Pilocarpinae (Rutaceae). Fl. Neotrop. Monogr. 33: 1-198. (Published by NYBG Press)
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Family
Rutaceae
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Scientific Name
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Type
Type. Palmer 175, Mexico. Guerrero: Acapulco, Dec 1895, fl, fr (lectotype, US-256910, the flowering part, photo F, NY; isolectotypes, DS, F 3x, GH, K, MO, NY, UC, US-256908, photo K, NY).
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Synonyms
Esenbeckia acapulcensis Rose
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Description
Subspecies Description - Indument of hairs 0.1-0.2 mm long; branchlets appressed-puberulous, becoming glabrous. Leaves 3-foliolate; petiole terete, semiterete and canaliculate towards the tip; leaflet blades with long-attenuate base to ca. 1 cm long, glabrous on both sides but ± appressed-pubescent at base, or quite appressed-pubescent particularly below. Partial inflorescences (5)10-13 x 4-10 cm, appressed-pubescent to somewhat tomentulose, often hoary, the branchlets tomentulose to minutely velvety towards the tips; bracts minutely pubescent distally above, appressed-puberulous below or (sub)glabrous on both sides. Flowers 5.5-7 mm in diam.; calyx lobes 0.8-1 × 1.2-1.5 mm, glabrous; petals deciduous, glabrous, venation imperfectly acrodromous but hardly visible except for the median nerve; filaments deciduous; carpels apically provided with apophyses 0.6-0.7 × 0.5-0.6 mm which are glabrous and very densely charged with roundish glandular tubercles, the longest tubercles occurring medio-dorsally on the loculi; the tubercles enlarging immediately after flowering and developing hyaline, spreading hairs to 0.3 mm long, which gradually disappear; style 1.1-1.4 mm, free for 0.5-0.6 mm. Fruits subglobose, depressed and stellately-lobed, ca. 2-3.2 × 2.7-3.5 cm when closed, when dehisced 1.7-2 × 3.3-4.5 cm, with 5 dorso-apical, erect, obtuse hornlike apophyses of 4-15 mm long, loculi medio-dorsally strongly carinate, irregularly (shortly) tuberculate-muricate, the tubercles to 5 mm; seed with a shortly curved beak at apex, chalazal area not visible.
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Discussion
Differences of the infrequently collected subsp. acapulcensis with the other subspecies are found in the smaller flowers and shorter style, and particularly in the fruits with the long dorso-apical horns. I cannot agree with Standley (1923) who considered it as synonymous with E. berlandieri (pro sp).
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Distribution
Mexico, allopatric with the other subspecies.
Mexico North America| Guerrero Mexico North America| Colima Mexico North America| Jalisco Mexico North America| Chihuahua Mexico North America|