Taxon Profile:

	The Plant List
	International Plant Name Index
	Tropicos
	Catalogue of Life
	Global Biodiversity Information Facility
	JSTOR Types
	JSTOR
	BHL
	Encyclopedia of Life
	WikiSpecies
	Google Scholar
	PubMed
	Morphbank
	IUCN
	National Center for Biotechnology Information
	Barcode of Life

Family:

Ericaceae (Magnoliophyta)

Primary Citation:

Bot. Jahrb. Syst. 42: 269. 1909

Accepted Name:

This name is currently accepted.

Common Names:

joyapa

Description:

Description: Terrestrial shrub to 3 m tall, from lignotuber to 2 m diam.; stem terete to subterete, strongly bluntly ribbed, nitid, slightly puberulous, glabrate; twigs subterete, complanate to bluntly or sharply angled, striate, densely short-pilose, glabrate. Leaves coriaceous, elliptic, ovate-elliptic, to rarely obovate, (2.3-)3-6.5(-9.5) x (1-)1.5-2.7(-4) cm, base obtuse to cuneate, apex obtuse or subacute, margin entire or faintly crenulate and sometimes slightly revolute near base, glabrous on both surfaces but deciduously minutely glandular-fimbriate beneath; pinnately nerved with 2-3 lateral nerves per side, midrib impressed above and raised beneath, lateral nerves arcuate-ascending, slightly impressed or plane above and slightly raised beneath, reticulate veinlets obscure; petiole subterete, flattened above, narrowly winged, rugose, 3-6(-8) mm long. Inflorescence subfasciculate, 2-4-flowered, surrounded at base by a few, triangular, puberulous, ciliate bracts ca. 2 mm long; floral bract sometimes coriaceous and dark in color at base becoming membranous and a lighter color distally, ovate, acute to acuminate, ca. 3-4 x 2.5-3 mm, dorsally short-pilose, glabrate; pedicel subterete, ribbed, 3-8(-10) mm long, densely short-pilose with pale spreading hairs ca. 0.2 mm long; bracteoles basal to medial, similar to floral bract but ca. 3(-4) mm long, usually with 1-2 circular, concave, blackish glands ca. 0.2-0.3 mm diam. at the base on either side of the midrib. Flowers with calyx 7-12.5 mm long, obscurely articulate with pedicel, short-pilose; hypanthium obprismatic, 10-ribbed to sharply 5(-6)-angled (narrowly winged), 2-3.5 mm long and 3 mm diam.; limb spreading-erect, 5-9 mm long; lobes 5(-6), striate to ribbed, lanceolate, acute-acuminate, 4.5-9 x 2-2.5 mm, marginally deciduously glandular-fimbriate; sinuses acute; corolla cylindric, 5(-6)-angled, 15-20 mm long and 4-6 mm diam., densely short-pilose, pale pink to red, the lobes spreading, oblong-ovate, bluntly acute, 1.5-3.5 mm long; stamen 10(-12), 7.5-10.5 mm long; filaments distinct, 0.5-3 mm long, glabrous; anthers ca. 9 mm long; thecae 3.5-5.5 mm long; tubules 2, laterally connate to tip, rarely the tubules completely fusing to 1 (and then dehiscing through one cleft), ca. 3-3.5 mm long, dehiscing by clefts to 2 mm long; style about as long as corolla or exserted, glabrous. Mature berry not seen, but immature at least 15 mm diam., glabrous.

Distribution: Ecuador and Peru; montane forest, southern Ecuadorean scrub vegetation, roadside slopes, at (1500-)2160-3400 m altitude.

Type: Ecuador. Loja: Cordillera near Loja, Jul-Aug (fl), Hartweg 784 (holotype, K; isotypes, B?, BM, BREM, CGE, E, K, L fragment, NY, OXF, P, W; photo of NY type NY neg. 9993).

Local name: Ecuador: joyapa

Cultivated: E.

Flora and Monograph Treatment(s):

Macleania salapa (Benth.) Hook.f. ex Hoerold: [Article] 1909. Bot. Jahrb. Syst. 42: 269.
Macleania salapa (Benth.) Hook.f. ex Hoerold: [Article] Smith, Albert C. 1952. Plants collected in Ecuador by W. H. camp. Vaceiniaceae. Mem. New York Bot. Gard. 8 (1): 41-85.