Taxon Details: Ceratostema nodosum Luteyn
Taxon Profile:
Narratives:
Family:
Ericaceae (Magnoliophyta)
Ericaceae (Magnoliophyta)
Scientific Name:
Ceratostema nodosum Luteyn
Ceratostema nodosum Luteyn
Accepted Name:
This name is currently accepted.
This name is currently accepted.
Description:
Description: Terrestrial shrub 1-2.5 m tall; stem and twigs subterete, bluntly angled, densely white-pilose with unicellular hairs to 1.3 mm long. Leaves oblong, ovate, or elliptic, (4-)7-15 x (2.5-)3-7(-8) cm, coriaceous, apex acute or short-acuminate, rarely obtuse or rounded, base deeply cordate, auriculate with lobes well separated, amplexicaul, margin entire, upper surface short-pilose along veins proximally, lower surface short-pilose throughout; pinnately nerved with 3-4 lateral nerves per side, these red when fresh, midrib and lateral nerves impressed above and raised beneath, veinlets raised on both surfaces; petiole 2-5 mm long, terete, rugose, densely white-pilose. Inflorescence solitary, axillary, racemose, ca. 15-20-flowered, nodding at anthesis but erect afterward; rachis subterete, bluntly angled, 2-4(-5.5) cm long, densely silky-pilose; floral bract membranous, lanceolate to oblanceolate, apically abruptly short-acuminate and somewhat mucronate, venation prominent, 40-70 x 20-35 mm, short-pilose, lower ones pale green to rose-red when fresh, upper ones rose-red; pedicel subterete, 10-12 mm long and 4 mm diam., densely silky-pilose; bracteoles located near or above middle of pedicel, basally imbricate, lanceolate, apically acute or acuminate, 30-45 mm long, short-pilose on both surfaces. Flowers with calyx articulate, 11-14 mm long, pale green, densely short-pilose; hypanthium subcylindrical, ca. 6-7 mm long, 10-costate; limb campanulate, 5-7 mm long; lobes broadly triangular, acute to acuminate, 2-3.5 mm long; sinuses broadly rounded; corolla cylindrical, terete, 41-45 mm long and ca. 8 mm diam., red, glabrous except short-pilose along lobes, the lobes narrowly triangular, long-acuminate, 12-16 mm long, dark purple within; stamen 10, alternately slightly unequal, 39-42 mm long; filaments about equal, distinct, 8.5-10 mm long, glabrous; anthers alternately slightly unequal, 31-35 mm long; thecae alternately slightly unequal, 11-15 mm long; tubules distinct to base where papillate, about equal, longer than thecae, 20.5-21.5 mm long, appearing to arise from dorsal side of thecae, dehiscing by subterminal, oblique clefts 1-1.5 mm long; nectariferous disc cupular, glabrous; style exserted, ca. 46-49 mm long, glabrous. Berry not seen.
Distribution: Endemic to Ecuador; premontane moist to rainforest to montane cloud forest at 1100-3000 m altitude.
Type: Ecuador. Napo: Baeza-Tena rd, 25-28 km S of bridge in Cosanga, 1375-1460 m, 10 Jan 1979 (fl), Luteyn & Lebrón-Luteyn 6745 (holotype, NY; isotypes, AAU, CAS, E, F, GB, GH, L, MO, P, QCA, US).
Description: Terrestrial shrub 1-2.5 m tall; stem and twigs subterete, bluntly angled, densely white-pilose with unicellular hairs to 1.3 mm long. Leaves oblong, ovate, or elliptic, (4-)7-15 x (2.5-)3-7(-8) cm, coriaceous, apex acute or short-acuminate, rarely obtuse or rounded, base deeply cordate, auriculate with lobes well separated, amplexicaul, margin entire, upper surface short-pilose along veins proximally, lower surface short-pilose throughout; pinnately nerved with 3-4 lateral nerves per side, these red when fresh, midrib and lateral nerves impressed above and raised beneath, veinlets raised on both surfaces; petiole 2-5 mm long, terete, rugose, densely white-pilose. Inflorescence solitary, axillary, racemose, ca. 15-20-flowered, nodding at anthesis but erect afterward; rachis subterete, bluntly angled, 2-4(-5.5) cm long, densely silky-pilose; floral bract membranous, lanceolate to oblanceolate, apically abruptly short-acuminate and somewhat mucronate, venation prominent, 40-70 x 20-35 mm, short-pilose, lower ones pale green to rose-red when fresh, upper ones rose-red; pedicel subterete, 10-12 mm long and 4 mm diam., densely silky-pilose; bracteoles located near or above middle of pedicel, basally imbricate, lanceolate, apically acute or acuminate, 30-45 mm long, short-pilose on both surfaces. Flowers with calyx articulate, 11-14 mm long, pale green, densely short-pilose; hypanthium subcylindrical, ca. 6-7 mm long, 10-costate; limb campanulate, 5-7 mm long; lobes broadly triangular, acute to acuminate, 2-3.5 mm long; sinuses broadly rounded; corolla cylindrical, terete, 41-45 mm long and ca. 8 mm diam., red, glabrous except short-pilose along lobes, the lobes narrowly triangular, long-acuminate, 12-16 mm long, dark purple within; stamen 10, alternately slightly unequal, 39-42 mm long; filaments about equal, distinct, 8.5-10 mm long, glabrous; anthers alternately slightly unequal, 31-35 mm long; thecae alternately slightly unequal, 11-15 mm long; tubules distinct to base where papillate, about equal, longer than thecae, 20.5-21.5 mm long, appearing to arise from dorsal side of thecae, dehiscing by subterminal, oblique clefts 1-1.5 mm long; nectariferous disc cupular, glabrous; style exserted, ca. 46-49 mm long, glabrous. Berry not seen.
Distribution: Endemic to Ecuador; premontane moist to rainforest to montane cloud forest at 1100-3000 m altitude.
Type: Ecuador. Napo: Baeza-Tena rd, 25-28 km S of bridge in Cosanga, 1375-1460 m, 10 Jan 1979 (fl), Luteyn & Lebrón-Luteyn 6745 (holotype, NY; isotypes, AAU, CAS, E, F, GB, GH, L, MO, P, QCA, US).
Flora and Monograph Treatment(s):
Ceratostema nodosum Luteyn: [Article] 1986. J. Arnold Arbor. 67: 489.
Ceratostema nodosum Luteyn: [Article] 1986. J. Arnold Arbor. 67: 489.
Related Objects:
• D. A. Neill 5935, Ecuador
• D. A. Neill 8378, Ecuador
• E. Asplund 9843, Ecuador
• J. L. Luteyn 15045, Ecuador
• J. L. Luteyn 15347, Ecuador
• J. L. Luteyn 15347, Ecuador
• J. L. Luteyn 14662, Ecuador
• J. L. Luteyn 14662, Ecuador
• J. L. Luteyn 6739, Ecuador
• J. L. Luteyn 14691, Ecuador
• J. L. Luteyn 14652, Ecuador
• J. L. Luteyn 13457, Ecuador
• J. D. Boeke 941, Ecuador
• C. E. Cerón Martínez 2996, Ecuador
• C. E. Cerón Martínez 2957, Ecuador
• J. L. Luteyn 6745, holotype; South America
• J. L. Luteyn 6745, holotype; South America
• Collector unknown 95, Ecuador
• C. E. Cerón Martínez 6527, Ecuador
• F. Hurtado 2594, Ecuador
• W. A. Palacios 9540, Ecuador
• D. A. Neill 10984, Ecuador
• D. A. Neill 8378, Ecuador
• E. Asplund 9843, Ecuador
• J. L. Luteyn 15045, Ecuador
• J. L. Luteyn 15347, Ecuador
• J. L. Luteyn 15347, Ecuador
• J. L. Luteyn 14662, Ecuador
• J. L. Luteyn 14662, Ecuador
• J. L. Luteyn 6739, Ecuador
• J. L. Luteyn 14691, Ecuador
• J. L. Luteyn 14652, Ecuador
• J. L. Luteyn 13457, Ecuador
• J. D. Boeke 941, Ecuador
• C. E. Cerón Martínez 2996, Ecuador
• C. E. Cerón Martínez 2957, Ecuador
• J. L. Luteyn 6745, holotype; South America
• J. L. Luteyn 6745, holotype; South America
• Collector unknown 95, Ecuador
• C. E. Cerón Martínez 6527, Ecuador
• F. Hurtado 2594, Ecuador
• W. A. Palacios 9540, Ecuador
• D. A. Neill 10984, Ecuador