Monographs Details:
Authority:
Luteyn, James L. 1983. Ericaceae--part I. Cavendishia. Fl. Neotrop. Monogr. 35: 1-290. (Published by NYBG Press)
Luteyn, James L. 1983. Ericaceae--part I. Cavendishia. Fl. Neotrop. Monogr. 35: 1-290. (Published by NYBG Press)
Family:
Ericaceae
Ericaceae
Description:
Species Description - Epiphytic shrub; branches subterete or bluntly angled, slightly striate, glabrous, with scattered pustular glands to 0.3 mm diam., reddish-brown. Leaves lanceolate or elliptic, (5-)7-14.5 X 2-5.5 cm, basally obtuse or narrowly rounded, apically acuminate (sometimes abruptly short caudate-acuminate), glabrous but with scattered black sessile and pustular glands 0.1 mm in diam. along upper leaf surface, the sessile ones often caducous leaving a reddish punctate scar; 5(-7)-plinerved, innermost lateral nerves arising to 1.5 cm above base, midrib and lateral nerves deeply impressed above and conspicuously raised beneath causing leaves to be bullate, veinlets slightly impressed above and raised beneath; petioles subterete, rugose, 3-5 mm long and 2.5-3.5 mm in diam., glabrous or hispid only on abaxial surface, provided with globular glands 0.2 mm in diam. in distal portion. Inflorescence viscid, ca. 20-flowered but lowest few nodes sterile; rachis subterete, bluntly angled, striate, glabrous, 11-14 cm long (upper portion still in bud) and 3-4 mm in diam. with scattered globular or angular glands ca. 0.2 mm long; floral bracts ovate, oblong or oval-elliptic, 21-25 X 10-12 mm, glabrous or weakly ciliate, basally narrowed, truncate and clasping, apically rounded, marginally crisped and with 12-20 dark red globular glands 0.1-0.2 mm in diam. on each side, pink; pedicels subterete, swollen distally, striate, glabrous, 5-7 mm long and 0.75 mm in diam., with globular glands concentrated at distal swollen portion; bracteoles oblong-oblanceolate, 10-14 x 6-8 mm, located midway along pedicel and clasping or nearly concealing entire calyx and lower 1/5 of corolla at anthesis, margins crisped and with dark red globular glands scattered along edge. Flowers: calyx glabrous, ca. 3.5-5 mm long; hypanthium cylindric, obscurely ribbed, 1.5-2 mm long, truncate basally, covered by globular or angular glands 0.2 mm in diam.; limb spreading-campanulate, 2-3 mm long and covered by globular glands; lobes broadly triangular, 1 mm long, erect after anthesis, completely glandular-callose distal 2/3 and covered by globular glands; sinuses flat to broadly rounded; corolla slightly narrowed distally, glabrous without but sparsely pilose within, ca. 19 mm long and 4 mm in diam., pink when fresh, lobes triangular, acute or obtuse, 1 mm long; stamens 17 mm long; filaments alternately ca. 2.5 mm and 6 mm long, the short ones sparsely pilose dorsally at distal tips, the long ones densely pilose ventrally in distal half; anthers alternately ca. 16.5 mm and 13 mm long; thecae ca. 7 mm long; style ca. 19 mm long. Berry not seen.
Species Description - Epiphytic shrub; branches subterete or bluntly angled, slightly striate, glabrous, with scattered pustular glands to 0.3 mm diam., reddish-brown. Leaves lanceolate or elliptic, (5-)7-14.5 X 2-5.5 cm, basally obtuse or narrowly rounded, apically acuminate (sometimes abruptly short caudate-acuminate), glabrous but with scattered black sessile and pustular glands 0.1 mm in diam. along upper leaf surface, the sessile ones often caducous leaving a reddish punctate scar; 5(-7)-plinerved, innermost lateral nerves arising to 1.5 cm above base, midrib and lateral nerves deeply impressed above and conspicuously raised beneath causing leaves to be bullate, veinlets slightly impressed above and raised beneath; petioles subterete, rugose, 3-5 mm long and 2.5-3.5 mm in diam., glabrous or hispid only on abaxial surface, provided with globular glands 0.2 mm in diam. in distal portion. Inflorescence viscid, ca. 20-flowered but lowest few nodes sterile; rachis subterete, bluntly angled, striate, glabrous, 11-14 cm long (upper portion still in bud) and 3-4 mm in diam. with scattered globular or angular glands ca. 0.2 mm long; floral bracts ovate, oblong or oval-elliptic, 21-25 X 10-12 mm, glabrous or weakly ciliate, basally narrowed, truncate and clasping, apically rounded, marginally crisped and with 12-20 dark red globular glands 0.1-0.2 mm in diam. on each side, pink; pedicels subterete, swollen distally, striate, glabrous, 5-7 mm long and 0.75 mm in diam., with globular glands concentrated at distal swollen portion; bracteoles oblong-oblanceolate, 10-14 x 6-8 mm, located midway along pedicel and clasping or nearly concealing entire calyx and lower 1/5 of corolla at anthesis, margins crisped and with dark red globular glands scattered along edge. Flowers: calyx glabrous, ca. 3.5-5 mm long; hypanthium cylindric, obscurely ribbed, 1.5-2 mm long, truncate basally, covered by globular or angular glands 0.2 mm in diam.; limb spreading-campanulate, 2-3 mm long and covered by globular glands; lobes broadly triangular, 1 mm long, erect after anthesis, completely glandular-callose distal 2/3 and covered by globular glands; sinuses flat to broadly rounded; corolla slightly narrowed distally, glabrous without but sparsely pilose within, ca. 19 mm long and 4 mm in diam., pink when fresh, lobes triangular, acute or obtuse, 1 mm long; stamens 17 mm long; filaments alternately ca. 2.5 mm and 6 mm long, the short ones sparsely pilose dorsally at distal tips, the long ones densely pilose ventrally in distal half; anthers alternately ca. 16.5 mm and 13 mm long; thecae ca. 7 mm long; style ca. 19 mm long. Berry not seen.
Discussion:
Cavendishia fortunensis is a typical member of ser. Lactiviscidae, although at this time I do not know if the inflorescence exudate is clear or milky-white. The species has a superficial similarity to C. albopicata and C. pseudostenophylla because of the medial position of the bracteoles along the pedicel. However, I feel it is more closely related to C. panamensis and C. arizonensis because of its overall coarser habit, nonapophysate calyx, and hypanthium which is nearly obscured from view by the abundance of globular to irregularly spherical glands. It also has in common with C. arizonensis bullate (not flat) leaves, and with C. panamensis globular-glandular (not glandular-callose) bracteoles. From these two species, C. fortunensis differs by the medial (not basal) insertion of the larger bracteoles (10-14 mm not 1.5-4 mm long), and glandular calyx limb.
Cavendishia fortunensis is a typical member of ser. Lactiviscidae, although at this time I do not know if the inflorescence exudate is clear or milky-white. The species has a superficial similarity to C. albopicata and C. pseudostenophylla because of the medial position of the bracteoles along the pedicel. However, I feel it is more closely related to C. panamensis and C. arizonensis because of its overall coarser habit, nonapophysate calyx, and hypanthium which is nearly obscured from view by the abundance of globular to irregularly spherical glands. It also has in common with C. arizonensis bullate (not flat) leaves, and with C. panamensis globular-glandular (not glandular-callose) bracteoles. From these two species, C. fortunensis differs by the medial (not basal) insertion of the larger bracteoles (10-14 mm not 1.5-4 mm long), and glandular calyx limb.
Distribution and Ecology: Cavendishia fortunensis is known only from the type locality where it flowers in April and May. Although the label (of the type) does not record habitat, the area around the Fortuna dam site ranges from ca. 1100- 1800 m and contains cloud forest.
Distribution:
Panama Central America| Chiriquí Panamá Central America|
Panama Central America| Chiriquí Panamá Central America|
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